How Irreducibly Complex Systems Evolve. Data Behe must deal with.

From: glenn morton (glenn.morton@btinternet.com)
Date: Sun Oct 15 2000 - 11:18:52 EDT

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    Marcio Pie pointed us to a fascinating article that up to this moment no one
    has responded to. The silence on the part of the Christian apologists, most
    noteably the ID folk who believe that irreducibly complex systems can't
    arise without divine help, seems rather odd. The article that Marcio pointed
    us to has gone a long way toward explaining how such systems can arise--yet
    the apologists ignore this article. IMO this is an extremely important
    article in the creation/evolution debate.

    Marcio wrote:
    > An interesting article came out recently in Nature on how a prion may
    > improve the "evolvability" of yeast. This may have some relevance
    > regarding some ID hypotheses. I'd love to hear your comments on this.

    And then cited the first paragraph:

    > Nature 407, 477 - 483 (2000)
    >
    > A yeast prion provides a mechanism for genetic variation and phenotypic
    > diversity
    >
    > HEATHER L. TRUE AND SUSAN L. LINDQUIST
    >
    > A major enigma in evolutionary biology is that new forms or functions
    > often require the concerted effects of several independent genetic
    > changes. It is unclear how such changes might accumulate when they are
    > likely to be deleterious individually and be lost by selective pressure.
    > The Saccharomyces cerevisiae prion [PSI+] is an epigenetic modifier of the
    > fidelity of translation termination, but its impact on yeast biology has
    > been unclear. Here we show that [PSI+] provides the means to uncover
    > hidden genetic variation and produce new heritable phenotypes. Moreover,
    > in each of the seven genetic backgrounds tested, the constellation of
    > phenotypes produced was unique. We propose that the epigenetic and
    > metastable nature of [PSI+] inheritance allows yeast cells to exploit
    > pre-existing genetic variation to thrive in fluctuating environments.
    > Further, the capacity of [PSI+] to convert previously neutral genetic
    > variation to a non-neutral state may facilitate the evolution of new
    > traits.
    >

    The second paragraph tells us a bit about how this happens:

            “Here we report that a protein-based element of inheritance (a prion) in
    the yeast Sacharomyces cerevisiae provides a means to expose this and other
    types of silent protein-coding information. The prion [PSI+] reduces the
    fidelity with which ribosomes terminate translation at stop codons in a
    metastable, heritable manner. The reduction in translational fidelity caused
    by [PSI+] is routinely monitored by the production of active products from
    genetic markers containing stop-codon mutations.” Heather L. True and Susan
    L. Lindquist, “A Yeast Prion Provides a Mechanism For Genetic Variation and
    Phenotypic Diversity,” Nature, 407(2000):477-483, p. 477

    This prion seems to make the protein manufacturing facility--the
    ribosome--pay less attention to the instructions to stop reading the gene.
    This means that the data on the genome past the gene will be read. It may be
    nonsense or it may be something good. In any event, the prion causes parts
    of the genome which are not normally sampled to be tested for useful
    codings. If useful proteins are found in these stranded parts of the genome,
    then they manufactured and put out into the cell. If several proteins (from
    different parts of the genome) are needed to work in concert in order for
    the cell to survive, the prion allows them to be found and used. The prion,
    by causing the stop codons to be ignored, allow a period of testing of many,
    many novel places in the genome.

     True and Lindquist then tell how this mechanism causes new evolution. They
    write:

            “We have demonstrated that the [PSI+] element of the yeast S. cerevisiae
    provides a means to unveil silent genetic information to produce new
    heritable phenotypes. IN the context of an individual cell, [PSI+] allows
    alternative heritable phenotypes to be encoded by a single genome. In the
    context of diverging populations it provides a vast array of new phenotypic
    states with unique growth advantages and disadvantages. We propose that the
    epigenetic and metastable nature of [PSI+], associated with the fundamental
    process of translation, potentiates survival in a fluctuating environment
    and provides a conduit for the evolution of new traits.
            “Yeast cells spontaneously switch from the [psi-] to the PSI+] state, with
    rates generally varying between 1 in 10^5 to 1 in 10^7. Thus, once a
    population has reached an appreciable size, some of its genetically
    identical members will have acquired a new, heritable phenotype. If the
    environment does not favour this phenotype, the loss of these cells will
    have a minimal impact on the fitness of that genotype. If the environment
    does favour it, the original genotype can thrive even if the original
    phenotype is disfavored. Once the progeny with the new phenotype have
    reached sufficient densities the spontaneous appearance of [psi-]
    derivatives will ensure that the original phenotype is available should the
    environment change.”
            “[PSI+] allows cells to occupy a new niche without foregoing their capacity
    to occupy the old. If the new phenotype remains advantageous, the size of
    the growing population increases the likelihood that mutations will arise to
    eliminate stop codons that are relevant to the phenotype and require [PSI+]
    for read-through. These would make that phenotype more robust and fix the
    trait; cells would retain the phenotype upon reversion to the [psi-] state
    (presumably normal translation fidelity will generally be favoured in
    nature). By this mechanism, [PSI+] could facilitate the evolution of new
    traits.” Heather L. True and Susan L. Lindquist, “A Yeast Prion Provides a
    Mechanism For Genetic Variation and Phenotypic Diversity,” Nature,
    407(2000):477-483, p. 481- 482

    Thus, if the environment requires these novel proteins in order to survive,
    cells without them will die but cells with them and with the positive prion
    state will live. Eventually mutation will remove the stop codons which
    normally prevent the manufacture of these novel proteins.

    This mechanism answers has huge implications for Behe's claim. Michael
    writes:

            "Well, for starters, a system that is irreducibly complex. By irreducibly
    complex I mean a single system composed of several well-matched, interacting
    parts that contribute to the basic function, wherein the removal of any one
    of the parts causes the system to effectively cease functioning. An
    irreducibly complex system cannot be produced directly (that is, by
    continuously improving the initial function, which continues to work by the
    same mechanism) by slight, successive modifications of a precursor system
    because any precursor to an irreducibly complex system that is missing a
    part is by definition nonfunctional." ~ Michael J. Behe, Darwin's Black Box,
    (New York: The Free Press, 1996), p. 39

    Even if Behe is correct, that such systems can't arise by small independent
    changes, this mechanism allows for such systems to arise all at once by
    natural causes. Thus, irreducibly complex systems can't be considered
    indicative of designers--either intelligent alien life forms or divine
    beings. This mechanism effectively refutes the use of complex systems to
    infer design.

    Behe can no longer claim that as the number of working components increase,
    the likelihood of a natural route to the system goes down. Behe claimed,

            "Even if a system is irreducibly complex (and thus cannot have been
    produced directly), however, one can not definitively rule out the
    possibility of an indirect, circuitous route. As the complexity of an
    interacting system increases, though, the likelihood of such an indirect
    route drops precipitously. And as the number of unexplained, irreducibly
    complex biological systems increases, our confidence that Darwin's criterion
    of failure has been met skyrockets toward the maximum that science allows."
    ~ Michael J. Behe, Darwin's Black Box, (New York: The Free Press, 1996), p.
    40

    The prion system allows lots of different places on the genome to be tested
    for useful codings. And given the fact (that I pointed out earlier) that
    useful codings are found at a rate of 10^-13 in randomly made sequences, one
    must realize that such a mechanism will allow for this search process to be
    successful!

    The ID group needs to cease looking for evidence of design in biological
    systems. It isn't there. While I believe that the universe is designed, I
    simply don't think that biological systems are capable of yielding evidence
    for design. As such, we should be honest with ourselves and our readers.

    Thank you Marcio for pointing out this article and its importance. Why have
    the anti-evolutionary apologists not pointed it out first?

    glenn

    see http://www.flash.net/~mortongr/dmd.htm
    for lots of creation/evolution information

    >



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